HR: 12:05h
AN: B21G-08 [PDF]
TI: Calcite and Picocyanobacteria in Lakes: Factors Affecting Their Interaction
AU: * Dittrich, M
EM: dittrich@eawag.ch
AF: Swiss Federal Institute for Environmental Science and Technology, EAWAG, Limnological Research Center,
Seestrasse 79, Kastanienbaum, 6047
Switzerland
AU: Obst, M
AF: Swiss Federal Institute for Environmental Science and Technology, EAWAG, Limnological Research Center,
Seestrasse 79, Kastanienbaum, 6047
Switzerland
AU: Mavrocordatos, D
AF: Swiss Federal Institute for Environmental Science and Technology, EAWAG, Ueberlandstrasse 133,
Duebendorf, 8600
Switzerland
AB:
Calcites build large deposits which have been observed in the rock record throughout geological time at various localities
around the globe. Carbonate deposits have affected atmospheric carbon dioxide concentration. As it has been generally
accepted, inorganic precipitation represents a source of carbon dioxide on short geological time scales and a sink of
inorganic carbon at long time scales from millions to thousands of millions years. However, recent research indicates that
calcite deposits may result from microbial calcification instead of inorganic precipitation. In this case the process may
reduce atmospheric carbon dioxide on geologically short time scales. Thus the effect of carbonate sediment deposition on
global carbon cycling depends on the origin of carbonate. Thus it is essential to understand the cause and the key parameters
affecting calcite precipitation.
The role of algae and bacteria in calcite formation in lakes has not been evaluated in detail. Some evidence, however, exists
supporting precipitation of calcium carbonate by microbes as the origin of whiting. Several field studies on lakes have also
produced puzzling results: The peaks of algal blooms were often not found at the same time as precipitation events of
calcite.
We suspect that parts of the discrepancies in the interpretation of field observations are due to the activity of autotrophic
picoplankton. The unicellular autotrophic picoplankton (APP) is a ubiquitous component of pelagic ecosystems. But it has
often been overlooked due to its small cell size of 0.2 - 2 $\mu$m in diameter. Coccoid picocyanobacteria of the
Synechococcus-type dominate the picoplankton community in most oligotrophic systems. Recently, laboratory experiments and
field observations suggested that APP may play an important role in calcite precipitation.
The aim of this study was to examine the influence of environmental factors such as saturation state, concentration of
different dissolved ions and characteristics of the surface of cells on interaction between calcite and picocyanobacteria
under both laboratory and field conditions.
Laboratory experiments were performed with a picocyanobacteria strain Synechococcus-type. Using ion selective electrodes we
monitored calcite precipitation induced by bacteria in the solutions of a different composition (calcium 0.7 - 48 mM,
inorganic carbonate 6 - 35 $\mu$M). Electron and atomic force microscopy measurements provided insight into the cell-mineral
interface. Furthermore, quantitative investigations of the types and densities of proton binding sites on a bacterial surface
will be reported from the acid-base titrations on bacteria.
Results of these initial experiments are encouraging and demonstrate by direct measurements the potential of
picocyanobacteria to precipitate calcite. The amount of the precipitated calcite varied in experiments with a different ratio
of dissolved inorganic carbon and calcium. The microscopic observations provide some evidence that the cell walls of
cyanobacteria act as a substrate of nucleation of calcite.
Temporal and spatial correlations of cyanobacteria and calcite, as well as images of bacterial shape particles indicated that
picoplankton plays an important role in calcite precipitation in Lake Lucerne. This class of phytoplankton has to be
considered in studying the biogeochemical cycling of oligotrophic hardwater lakes.
DE: 0400 Biogeosciences
DE: 4805 Biogeochemical cycles (1615)
SC: Biogeosciences [B]
MN: 2003 Fall Meeting